S. Renaud and V. Millien, Intra-and interspecific morphological variation in the field mouse species Apodemus argenteus and A. speciosus in the Japanese archipelago: the role of insular isolation and biogeographic gradients, Biological Journal of the Linnean Society, vol.74, pp.557-569, 2001.
URL : https://hal.archives-ouvertes.fr/hal-00698172

F. Rohlf and D. Slice, Extensions of the Procrustes method for the optimal superimposition of landmarks, Systematic Biology, vol.39, pp.40-59, 1999.

U. Salzmann and P. Hoelzmann, The Dahomey Gap: an abrupt climatically induced rain forest fragmentation in West Africa during the late Holocene, The Holocene, vol.15, pp.190-199, 2005.

S. Schlager, Morpho and Rvcg -shape analysis in R, Statistical shape and deformation analysis, pp.217-256, 2017.

B. Sidlauskas, J. Mol, and R. Vari, Dealing with allometry in linear and geometric morphometrics: a taxonomic case study in the Leporinus cylindriformis group (Characiformes: Anostomidae) with description of a new species from Suriname, Zoological Journal of the Linnean Society, vol.162, pp.103-130, 2011.

J. Sukumaran and L. L. Knowles, Multispecies coalescent delimits structure, not species, Proceedings of the National Academy of Sciences of the United States of America, vol.114, pp.1607-1612, 2017.

S. Sveegaard, A. Galatius, R. Dietz, L. Kyhn, J. C. Koblitz et al., Defining management units for cetaceans by combining genetics, morphology, acoustics and satellite tracking, Global Ecology and Conservation, vol.3, pp.839-850, 2015.

P. Vanzolini and E. Williams, The vanishing refuge: a mechanism for ecogeographic speciation, Papéis Avulsos de Zoologia, vol.34, pp.251-255, 1981.

W. N. Venables and B. D. Ripley, Modern applied statistics with S, 2002.

C. Villemant, G. Simbolotti, and M. Kenis, Discrimination of Eubazus (Hymenoptera, Braconidae) sibling species using geometric morphometrics analysis of wing venation, Systematic Entomology, vol.32, pp.625-634, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00173812

K. L. Voje, T. F. Hansen, C. K. Egset, G. H. Bolstad, and C. Pélabon, Allometric constraints and the evolution of allometry, Evolution, vol.68, pp.866-885, 2014.

V. Koenigswald and W. , Order Pholidota, The Miocene land mammals of Europe, pp.75-80, 1999.

H. Zhang, M. Miller, F. Yang, H. K. Chan, P. Gaubert et al., Molecular tracing of confiscated pangolin scales for conservation and illegal trade monitoring in Southeast Asia, Global Ecology and Conservation, vol.4, pp.414-422, 2015.

Z. M. Zhou, Y. Zhou, C. Newman, and D. W. Macdonald, Scaling up pangolin protection in China, Frontiers in Ecology and the Environment, vol.12, pp.97-98, 2014.

, SUPPORTING INFORMATION Additional supporting information may be found in the online version of this article at the publisher's web-site: Appendix S1. Specimen list and landmark coordinates

S. Al,

, The Linnean Society of London, Zoological Journal of the Linnean Society, vol.188, pp.255-275, 2020.