T. J. Smayda, Harmful algal blooms: Their ecophysiology and general relevance to phytoplankton blooms in the sea, Limnology and Oceanography, vol.42, issue.5part2, pp.1137-1153, 1997.
DOI : 10.4319/lo.1997.42.5_part_2.1137

A. Zingone, The diversity of harmful algal blooms: a challenge for science and management, Ocean & Coastal Management, vol.43, issue.8-9, pp.725-748, 2000.
DOI : 10.1016/S0964-5691(00)00056-9

P. M. Glibert, D. M. Anderson, P. Gentien, E. Granéeli, and K. G. Sellner, The Global, Complex Phenomena of Harmful Algal Blooms, Oceanography, vol.18, issue.2, pp.136-147, 2005.
DOI : 10.5670/oceanog.2005.49

S. E. Shumway, S. M. Allen, and P. Dee-boersma, Marine birds and harmful algal blooms: sporadic victims or under-reported events?, Harmful Algae, vol.2, issue.1, pp.1-17, 2003.
DOI : 10.1016/S1568-9883(03)00002-7

C. J. Bolch and M. F. De-salas, A review of the molecular evidence for ballast water introduction of the toxic dinoflagellates Gymnodinium catenatum and the Alexandrium ???tamarensis complex??? to Australasia, Harmful Algae, vol.6, issue.4, pp.465-485, 2007.
DOI : 10.1016/j.hal.2006.12.008

M. Laabir and P. Gentien, Survival of toxic dinoflagellates after gut passage in the pacific oyster Crassostrea gigas thunburg, J. Shellfish Res, vol.18, pp.217-222, 1999.

J. M. Burkholder, P. M. Glibert, and H. M. Skelton, Mixotrophy, a major mode of nutrition for harmful algal species in eutrophic waters, Harmful Algae, vol.8, issue.1, pp.77-93, 2008.
DOI : 10.1016/j.hal.2008.08.010

N. Takada, N. Umemura, K. Suenaga, T. Chou, A. Nagatsu et al., Pinnatoxins B and C, the most toxic components in the pinnatoxin series from the Okinawan bivalve Pinna muricata, Tetrahedron Letters, vol.42, issue.20, pp.3491-3494, 2001.
DOI : 10.1016/S0040-4039(01)00480-4

A. I. Selwood, C. O. Miles, A. L. Wilkins, R. Van-ginkel, R. Munday et al., Isolation, Structural Determination and Acute Toxicity of Pinnatoxins E, F and G, Journal of Agricultural and Food Chemistry, vol.58, issue.10, pp.6532-6542, 2010.
DOI : 10.1021/jf100267a

L. Rhodes, K. Smith, A. Selwood, P. Mcnabb, R. Van-ginkel et al., Production of pinnatoxins by a peridinoid dinoflagellate isolated from Northland, New Zealand, Harmful Algae, vol.9, issue.4, pp.384-389, 2010.
DOI : 10.1016/j.hal.2010.01.008

E. Nezan, N. Chomerat, and . Vulcanodinium, gen. et sp. nov. (Dinophyceae), un Nouveau Dinoflagell?? Marin de la C??te M??diterran??enne Fran??aise, Cryptogamie, Algologie, vol.32, issue.1, pp.3-18, 2011.
DOI : 10.7872/crya.v32.iss1.2011.003

P. Hess, E. Abadie, F. Herve, T. Berteaux, V. Sechet et al., Pinnatoxin G is responsible for atypical toxicity in mussels (Mytilus galloprovincialis) and clams (Venerupis decussata) from Ingril, a French Mediterranean lagoon, Toxicon, vol.75, pp.16-26, 2013.
DOI : 10.1016/j.toxicon.2013.05.001
URL : https://hal.archives-ouvertes.fr/hal-00919909

L. Rhodes, K. Smith, A. Selwood, P. Mcnabb, S. Molenaar et al., Production of pinnatoxins E, F and G by scrippsielloid dinoflagellates isolated from Franklin Harbour, South Australia, New Zealand Journal of Marine and Freshwater Research, vol.41, issue.4, pp.703-709, 2011.
DOI : 10.1021/jf100267a

M. J. Garrett, C. Puchulutegui, A. I. Selwood, and J. Wolny, Identification of the harmful dinoflagellate Vulcanodinium rugosum recovered from a ballast tank of a globally traveled ship in Port Tampa Bay, Florida, USA, Harmful Algae, vol.39, pp.202-209, 2014.
DOI : 10.1016/j.hal.2014.07.014

K. F. Smith, L. L. Rhodes, S. Suda, and A. I. Selwood, A dinoflagellate producer of pinnatoxin G, isolated from sub-tropical Japanese waters, Harmful Algae, vol.10, issue.6, pp.702-705, 2011.
DOI : 10.1016/j.hal.2011.05.006

D. U. Hernandez-becerril, M. C. Rodriguez-palacio, and C. Lozano-ramirez, Morphology and life stages of the potentially pinnatoxin-producing thecate dinoflagellate Vulcanodinium rugosum from the tropical Mexican Pacific, Botanica Marina, vol.56, issue.5-6, pp.535-540, 2013.
DOI : 10.1515/bot-2013-0079

A. I. Selwood, A. L. Wilkins, R. Munday, H. F. Gu, K. F. Smith et al., Rise, F. Pinnatoxin H: A new pinnatoxin analogue from a south China sea Vulcanodinium. rugosum isolate, Tetrahedron Lett, vol.55, pp.2014-5508

L. Rhodes, K. Smith, A. Selwood, P. Mcnabb, R. Munday et al., Dinoflagellate Vulcanodinium rugosum identified as the causative organism of pinnatoxins in Australia, New Zealand and Japan, Phycologia, vol.50, issue.6, pp.624-628, 2011.
DOI : 10.2216/11-19.1

D. M. Anderson, P. M. Glibert, and J. M. Burkholder, Harmful algal blooms and eutrophication: Nutrient sources, composition, and consequences, Estuaries, vol.11, issue.4, pp.704-726, 2002.
DOI : 10.1007/BF02804901

S. C. Leong, A. Murata, Y. Nagashima, and S. Taguchi, Variability in toxicity of the dinoflagellate Alexandrium tamarense in response to different nitrogen sources and concentrations, Toxicon, vol.43, issue.4, pp.407-415, 2004.
DOI : 10.1016/j.toxicon.2004.01.015

T. C. Lee, O. T. Kwok, K. C. Ho, and F. W. Lee, Effects of different nitrate and phosphate concentrations on the growth and toxin production of an Alexandrium tamarense strain collected from Drake Passage, Marine Environmental Research, vol.81, pp.62-69, 2012.
DOI : 10.1016/j.marenvres.2012.08.009

P. T. Lim, C. P. Leaw, A. Kobiyama, and T. Ogata, Growth and toxin production of tropical Alexandrium minutum Halim (Dinophyceae) under various nitrogen to phosphorus ratios, Journal of Applied Phycology, vol.47, issue.4
DOI : 10.1007/s10811-009-9443-8

J. Xu, A. Y. Ho, L. He, K. Yin, C. Hung et al., Effects of inorganic and organic nitrogen and phosphorus on the growth and toxicity of two Alexandrium species from Hong Kong, Harmful Algae, vol.16, pp.89-97, 2012.
DOI : 10.1016/j.hal.2012.02.006

R. I. Jones, Mixotrophy in planktonic protists as a spectrum of nutritional strategies, Mar. Microb. Food Webs, vol.8, pp.87-96, 1994.

R. I. Jones, Mixotrophy in planktonic protists: an overview, Freshwater Biology, vol.78, issue.2, pp.219-226, 2000.
DOI : 10.1046/j.1365-2427.2000.00672.x

K. Nygaard and A. Tobiesen, Bacterivory in algae: A survival strategy during nutrient limitation, Limnology and Oceanography, vol.38, issue.2, pp.273-279, 1993.
DOI : 10.4319/lo.1993.38.2.0273

H. J. Jeong, Y. D. Yoo, J. S. Kim, T. H. Kim, J. H. Kim et al., Mixotrophy in the Phototrophic Harmful Alga Cochlodinium polykrikoides (Dinophycean): Prey Species, the Effects of Prey Concentration, and Grazing Impact, The Journal of Eukaryotic Microbiology, vol.36, issue.5, pp.563-569, 2004.
DOI : 10.1046/j.1529-8817.2000.00009.x

R. M. Kudela, J. P. Ryan, M. D. Blakely, J. Q. Lane, and T. D. Peterson, Linking the physiology and ecology of Cochlodinium to better understand harmful algal bloom events: A comparative approach, Harmful Algae, vol.7, issue.3, pp.278-292, 2008.
DOI : 10.1016/j.hal.2007.12.016

Y. Collos, A. Vaquer, M. Laabir, E. Abadie, T. Laugier et al., Contribution of several nitrogen sources to growth of Alexandrium catenella during blooms in Thau lagoon, southern France, Harmful Algae, vol.6, issue.6, pp.781-789, 2007.
DOI : 10.1016/j.hal.2007.04.003

O. Mitamura and Y. Saijo, Decomposition of urea associated with photosynthesis of phytoplankton in coastal waters, Marine Biology, vol.10, issue.1, pp.67-72, 1975.
DOI : 10.1007/BF00393754

Y. Collos, B. Bec, C. Jauzein, E. Abadie, T. Laugier et al., Oligotrophication and emergence of picocyanobacteria and a toxic dinoflagellate in Thau lagoon, southern France, Journal of Sea Research, vol.61, issue.1-2, pp.61-68, 2009.
DOI : 10.1016/j.seares.2008.05.008

Y. Perrin and Y. Collos, Influence of temperature, salinity and irradiance on the growth and cell yield of the harmful red tide dinoflagellate Alexandrium catenella colonizing mediterranean waters, J. Plankton Res, vol.33, pp.1550-1563, 2011.

M. Laabir, N. Barre, J. Franco, C. Brunet, and E. Masseret, Collos, Y. Morphological, biochemical and growth characteristics of Alexandrium catenella (Whedon & Kofoid) Balech, a toxic dinoflagellate expanding in Mediterranean waters, Cah. Biol, vol.53, pp.365-372, 2012.

R. Pistocchi, L. Pezzolesi, F. Guerrini, S. Vanucci, C. Dell-'aversano et al., A review on the effects of environmental conditions on growth and toxin production of Ostreopsis ovata, Toxicon, vol.57, issue.3, pp.421-428, 2011.
DOI : 10.1016/j.toxicon.2010.09.013

F. Guerrini, L. Pezzolesi, A. Feller, M. Riccardi, P. Ciminiello et al., Comparative growth and toxin profile of cultured Ostreopsis ovata from the Tyrrhenian and Adriatic Seas, Toxicon, vol.55, issue.2-3, pp.211-220, 2010.
DOI : 10.1016/j.toxicon.2009.07.019

S. L. Morton, A. Vershinin, L. L. Smith, T. A. Leighfield, S. Pankov et al., Seasonality of Dinophysis spp. and Prorocentrum lima in Black Sea phytoplankton and associated shellfish toxicity, Harmful Algae, vol.8, issue.5, pp.629-636, 2009.
DOI : 10.1016/j.hal.2008.10.011

P. Vale, V. Veloso, and A. Amorim, Toxin composition of a Prorocentrum lima strain isolated from the Portuguese coast, Toxicon, vol.54, issue.2, pp.145-152, 2009.
DOI : 10.1016/j.toxicon.2009.03.026

Y. Collos, C. Gagne, M. Laabir, A. Vaquer, P. Cecchi et al., Nitrogenous nutrition of Alexandrium catenella (dinophyceae) in cultures and in Thau lagoon

Y. Collos, M. Lespilette, A. Vaquer, M. Laabir, and A. Pastoureaud, during nutrient pulses, African Journal of Marine Science, vol.28, issue.2, pp.313-318, 2006.
DOI : 10.2989/18142320609504169
URL : https://hal.archives-ouvertes.fr/hal-00153629

L. J. Twomey, M. F. Piehler, and H. W. Paerl, Phytoplankton uptake of ammonium, nitrate and urea in the Neuse River Estuary, NC, USA, Hydrobiologia, vol.31, issue.4, pp.123-134, 2005.
DOI : 10.1007/s10750-004-2403-z

D. M. Anderson, T. J. Alpermann, A. D. Cembella, Y. Collos, E. Masseret et al., The globally distributed genus Alexandrium: Multifaceted roles in marine ecosystems and impacts on human health, Harmful Algae, vol.14, pp.10-35, 2012.
DOI : 10.1016/j.hal.2011.10.012

E. H. John and K. J. Flynn, Growth dynamics and toxicity of Alexandrium fundyense (Dinophyceae): the effect of changing N???P supply ratios on internal toxin and nutrient levels, European Journal of Phycology, vol.35, issue.1, pp.11-23, 2000.
DOI : 10.1017/S0967026200002572

I. Hadjadji, E. Masseret, B. Plisson, M. Laabir, P. Cecchi et al., Clonal variation in physiological parameters of Alexandrium tamarense: Implications for biological invasions and maintenance, Cah. Biol, vol.53, pp.357-363, 2012.

P. J. Harrison, R. E. Waters, and F. J. Taylor, A BROAD SPECTRUM ARTIFICIAL SEA WATER MEDIUM FOR COASTAL AND OPEN OCEAN PHYTOPLANKTON, Journal of Phycology, vol.4, issue.1, pp.28-35, 1980.
DOI : 10.1007/BF00446694

J. Larsen, H. Kuosa, J. Ikavalko, and K. Kivi, A redescription of Scrippsiella hangoei (Schiller) comb. nov. ???a ???red tide??? dinoflagellate from the northern Baltic, Phycologia, vol.34, issue.2, pp.135-144, 1995.
DOI : 10.2216/i0031-8884-34-2-135.1

A. Kremp and M. W. Parrow, EVIDENCE FOR ASEXUAL RESTING CYSTS IN THE LIFE CYCLE OF THE MARINE PERIDINOID DINOFLAGELLATE, SCRIPPSIELLA HANGOEI1, Journal of Phycology, vol.28, issue.2, pp.400-409, 2006.
DOI : 10.1111/j.0022-3646.1979.00312.x

R. R. Guillard, Division rates In Handbook of Phycological Methods: Culture Methods and Growth Measurements, pp.289-311, 1973.

E. Alvarez, A. Lopez-urrutia, E. Nogueira, and S. Fraga, How to effectively sample the plankton size spectrum? A case study using FlowCAM, Journal of Plankton Research, vol.33, issue.7, pp.1119-1133, 2011.
DOI : 10.1093/plankt/fbr012

Y. Collos, F. Mornet, A. Sciandra, N. Waser, A. Larson et al., An optical method for the rapid measurement of micromolar concentrations of nitrate in marine phytoplankton cultures, Journal of Applied Phycology, vol.11, issue.2, pp.179-184, 1999.
DOI : 10.1023/A:1008046023487

F. Koroleff, K. Grasshoff, K. Kremling, and M. Ehrhardt, Determination of ammonium, In Method of Sea Water Analysis, pp.117-182, 1983.

A. Aminot and R. Kerouel, Dosage automatique de l'ur??e dans l'eau de mer: une m??thode tr??s sensible ?? la diac??tylmonoxime, Canadian Journal of Fisheries and Aquatic Sciences, vol.39, issue.1, pp.174-183, 1982.
DOI : 10.1139/f82-020

J. Neveux and F. Lantoine, Spectrofluorometric assay of chlorophylls and phaeopigments using the least squares approximation technique, Deep Sea Research Part I: Oceanographic Research Papers, vol.40, issue.9, pp.1747-1765, 1993.
DOI : 10.1016/0967-0637(93)90030-7