H. Pretzsch, Forest Dynamics, Growth and Yield, 2009.
DOI : 10.1007/978-3-540-88307-4

C. Sierra, H. Loescher, M. Harmon, A. Richardson, and D. Hollinger, Interannual variation of carbon fluxes from three contrasting evergreen forests: the role of forest dynamics and climate, Ecology, vol.13, issue.10, pp.2711-2723, 2009.
DOI : 10.1175/1520-0442(1995)008<0336:TSCIAI>2.0.CO;2

D. Newbery and M. Lingenfelder, Plurality of tree species responses to drought perturbation in Bornean tropical rain forest, Plant Ecology, vol.21, issue.1, pp.147-167, 2008.
DOI : 10.1007/s11258-008-9533-8

J. Rapp, M. Silman, J. Clark, C. Girardin, and D. Galiano, Intra- and interspecific tree growth across a long altitudinal gradient in the Peruvian Andes, Ecology, vol.93, issue.9, pp.2061-2072, 2012.
DOI : 10.1111/j.1365-2389.2009.01175.x

D. Sánchez-gomez, M. Zavala, D. Schalkwijk, I. Urbieta, and F. Valladares, Interversion de rangs dans la croissance des arbres en relation avec la taille de l???arbre, la comp??tition et les gradients climatiques pour quatre esp??ces foresti??res dominantes dans la canop??e, Annals of Forest Science, vol.244, issue.6, pp.605-605, 2008.
DOI : 10.1051/forest:2008040

L. Sack and P. Grubb, Why do species of woody seedlings change rank in relative growth rate between low and high irradiance?, Functional Ecology, vol.11, issue.2, pp.145-154, 2001.
DOI : 10.1139/cjfr-28-6-871

B. Hérault, B. Bachelot, L. Poorter, V. Rossi, and F. Bongers, Functional traits shape ontogenetic growth trajectories of rain forest tree species, Journal of Ecology, vol.39, issue.6, pp.1431-1440, 2011.
DOI : 10.1111/j.1365-2745.2011.01883.x

C. Antin, R. Pélissier, G. Vincent, and P. Couteron, Crown allometries are less responsive than stem allometry to tree size and habitat variations in an Indian monsoon forest, Trees, vol.92, issue.5, pp.1485-1495, 2013.
DOI : 10.1007/s00468-013-0896-7

L. Poorter and E. Arets, Light environment and tree strategies in a Bolivian tropical moist forest: an evaluation of the light partitioning hypothesis, Plant Ecology, vol.166, issue.2, pp.295-306, 2003.
DOI : 10.1023/A:1023295806147

O. Atkin, M. Schortemeyer, N. Mcfarlane, and J. Evans, Variation in the components of relative growth rate in 10 Acacia species from contrasting environments, Plant, Cell and Environment, vol.30, issue.10, pp.1007-1017, 1998.
DOI : 10.1086/337467

M. Westbeek, Analysis of differences in photosynthetic nitrogen use efficiency of alpine and lowland Poa species, Oecologia, vol.120, issue.1, pp.19-26, 1999.
DOI : 10.1007/s004420050828

H. Poorter, C. Remkes, and H. Lambers, Carbon and Nitrogen Economy of 24 Wild Species Differing in Relative Growth Rate, PLANT PHYSIOLOGY, vol.94, issue.2, pp.621-627, 1990.
DOI : 10.1104/pp.94.2.621

J. Grime, The role of plasticity in exploiting environmental heterogeneity. Exploitation of environmental heterogeneity by plants, pp.2-19, 1994.

F. Valladares and Ü. Niinemets, Shade Tolerance, a Key Plant Feature of Complex Nature and Consequences, Annual Review of Ecology, Evolution, and Systematics, vol.39, issue.1, pp.237-257, 2008.
DOI : 10.1146/annurev.ecolsys.39.110707.173506

P. Reich, The world-wide ???fast-slow??? plant economics spectrum: a traits manifesto, Journal of Ecology, vol.97, issue.2, pp.275-301, 2014.
DOI : 10.1111/1365-2745.12211

J. Denslow, Tropical Rainforest Gaps and Tree Species Diversity, Annual Review of Ecology and Systematics, vol.18, issue.1, pp.431-451, 1987.
DOI : 10.1146/annurev.es.18.110187.002243

J. Bloor and P. Grubb, Growth and mortality in high and low light: trends among 15 shade-tolerant tropical rain forest tree species, Journal of Ecology, vol.81, issue.1, pp.77-85, 2003.
DOI : 10.1046/j.1469-8137.1999.00425.x

M. Uriarte, J. Clark, J. Zimmerman, L. Comita, and J. Forero-montaña, Multidimensional trade-offs in species responses to disturbance: implications for diversity in a subtropical forest, Ecology, vol.93, issue.1, pp.191-205, 2012.
DOI : 10.1017/S0266467406003890

J. Clark, Individuals and the Variation Needed for High Species Diversity in Forest Trees, Science, vol.327, issue.5969, pp.1129-1132, 2010.
DOI : 10.1126/science.1183506

S. Mcmahon, C. Metcalf, and C. Woodall, High-Dimensional Coexistence of Temperate Tree Species: Functional Traits, Demographic Rates, Life-History Stages, and Their Physical Context, PLoS ONE, vol.65, issue.5, p.21305020, 2011.
DOI : 10.1371/journal.pone.0016253.s002

C. Violle, B. Enquist, B. Mcgill, L. Jiang, and C. Albert, The return of the variance: intraspecific variability in community ecology, Trends in Ecology & Evolution, vol.27, issue.4, pp.244-243, 2012.
DOI : 10.1016/j.tree.2011.11.014

URL : https://hal.archives-ouvertes.fr/hal-00739064

J. Clark, M. Dietze, S. Chakraborty, P. Agarwal, and I. Ibanez, Resolving the biodiversity paradox, Ecology Letters, vol.42, issue.8, pp.647-659, 2007.
DOI : 10.1890/02-4045

J. Lichstein, J. Dushoff, S. Levin, and S. Pacala, Intraspecific Variation and Species Coexistence, The American Naturalist, vol.170, issue.6, pp.807-818, 2007.
DOI : 10.1086/522937

B. Courbaud, G. Vieilledent, and G. Kunstler, Intra-specific variability and the competition???colonisation trade-off: coexistence, abundance and stability patterns, Theoretical Ecology, vol.89, issue.3, pp.61-71, 2012.
DOI : 10.1007/s12080-010-0095-8

N. Rüger, U. Berger, S. Hubbell, G. Vieilledent, and R. Condit, Growth Strategies of Tropical Tree Species: Disentangling Light and Size Effects, PLoS ONE, vol.91, issue.9, p.21966498, 2011.
DOI : 10.1371/journal.pone.0025330.s005

S. Gourlet-fleury, L. Blanc, N. Picard, P. Sist, and J. Dick, Grouping species for predicting mixed tropical forest dynamics: looking for a strategy, Annals of Forest Science, vol.62, issue.8, pp.785-796, 2005.
DOI : 10.1051/forest:2005084

URL : https://hal.archives-ouvertes.fr/hal-01032060

M. Swaine and T. Whitmore, On the definition of ecological species groups in tropical rain forests, Vegetatio, vol.103, issue.1-2, pp.81-86, 1988.
DOI : 10.1007/BF00044629

D. Ouédraogo, F. Mortier, S. Gourlet-fleury, V. Freycon, and N. Picard, Slow-growing species cope best with drought: evidence from long-term measurements in a tropical semi-deciduous moist forest of Central Africa, Journal of Ecology, vol.68, issue.6, pp.1459-1470, 2013.
DOI : 10.1111/1365-2745.12165

A. Zuur, E. Ieno, N. Walker, A. Savaliev, and G. Smith, Mixed effects models and extensions in ecology with R, 2009.
DOI : 10.1007/978-0-387-87458-6

R. Pélissier, J. Pascal, N. Ayyappan, B. Ramesh, and S. Aravajy, Tree demography in an undisturbed Dipterocarp permanent sample plot at Uppangala, Western Ghats of India, Ecology, vol.92, issue.6, p.17, 2011.
DOI : 10.1890/10-1991.1

J. Pascal and R. Pélissier, Structure and floristic composition of a tropical evergreen forest in southwest India, J Trop Ecol, vol.12, pp.195-218, 1996.

A. Cnaan, N. Laird, and P. Slasor, Using the general linear mixed model to analyse unbalanced repeated measures and longitudinal data, Statistics in Medicine, vol.16, issue.20, pp.2349-2380, 1997.
DOI : 10.1002/(SICI)1097-0258(19971030)16:20<2349::AID-SIM667>3.0.CO;2-E

K. Burnham and D. Anderson, Model selection and multimodel inference, 2002.
DOI : 10.1007/b97636

H. Akaike, A new look at the statistical model identification, IEEE Trans Autom Control, vol.19, 1974.

S. Rai, Diameter/height, and diameter/girth relationship of some rain forest tree species of Karnataka?India, Malays For, vol.42, pp.53-58, 1979.

P. West, Use of diameter increment and basal area increment in tree growth studies, Canadian Journal of Forest Research, vol.10, issue.1, pp.71-77, 1980.
DOI : 10.1139/x80-012

F. Sterck and F. Bongers, Ontogenetic Changes in Size, Allometry, and Mechanical Design of Tropical Rain Forest Trees, American Journal of Botany, vol.85, issue.2, pp.266-266, 1998.
DOI : 10.2307/2446315

A. Weiskittel, S. Garber, G. Johnson, D. Maguire, and R. Monserud, Annualized diameter and height growth equations for Pacific Northwest plantation-grown Douglas-fir, western hemlock, and red alder, Forest Ecology and Management, vol.250, issue.3, pp.266-278, 2007.
DOI : 10.1016/j.foreco.2007.05.026

G. Steneker and J. Jarvis, A Preliminary Study to Assess Competition in a White Spruce - Trembling Aspen Stand, The Forestry Chronicle, vol.39, issue.3, pp.334-336, 1963.
DOI : 10.5558/tfc39334-3

M. Moravie, M. Durand, and F. Houllier, Ecological meaning and predictive ability of social status, vigour and competition indices in a tropical rain forest (India), Forest Ecology and Management, vol.117, issue.1-3, pp.221-240, 1999.
DOI : 10.1016/S0378-1127(98)00480-0

C. Canham, P. Lepage, and K. Coates, A neighborhood analysis of canopy tree competition: effects of shading versus crowding, Canadian Journal of Forest Research, vol.34, issue.4, pp.778-787, 2004.
DOI : 10.1139/x03-232

M. Contreras, D. Affleck, and W. Chung, Evaluating tree competition indices as predictors of basal area increment in western Montana forests, Forest Ecology and Management, vol.262, issue.11, pp.1939-1949, 2011.
DOI : 10.1016/j.foreco.2011.08.031

L. Comita, H. Muller-landau, S. Aguilar, and S. Hubbell, Asymmetric Density Dependence Shapes Species Abundances in a Tropical Tree Community, Science, vol.329, issue.5989, pp.330-332, 2010.
DOI : 10.1126/science.1190772

T. Kohyama and T. Takada, One-sided competition for light promotes coexistence of forest trees that share the same adult height, Journal of Ecology, vol.91, issue.6, pp.1-11, 2012.
DOI : 10.1111/j.1365-2745.2012.02029.x

R. Pélissier, Tree spatial patterns in three contrasting plots of a southern Indian tropical moist evergreen forest, Journal of Tropical Ecology, vol.14, issue.01, pp.1-16, 1998.
DOI : 10.1017/S0266467498000017

A. Robert and M. Moravie, Topographic variation and stand heterogeneity in a wet evergreen forest of India, Journal of Tropical Ecology, vol.19, issue.06, pp.697-707, 2003.
DOI : 10.1017/S0266467403006096

J. Clark, Fire and Climate Change During the Last 750 Yr in Northwestern Minnesota, Ecological Monographs, vol.60, issue.2, pp.135-159, 1990.
DOI : 10.2307/1943042

D. Bates, M. Maechler, and B. Bolker, lme4: Linear mixed-effects models using S4 classes. R package version 0, pp.999999-1000001, 2013.

R. Team, R: A language and environment for statistical computing, Development Core. R, 2012.

D. Alder, F. Oavika, M. Sanchez, J. Silva, and P. Van-der-hout, A comparison of species growth rates from four moist tropical forest regions using increment-size ordination, International Forestry Review, vol.4, issue.3, pp.196-206, 2002.
DOI : 10.1505/IFOR.4.3.196.17398

S. Nakagawa and H. Schielzeth, from generalized linear mixed-effects models, Methods in Ecology and Evolution, vol.22, issue.2, pp.133-142, 2013.
DOI : 10.1111/j.2041-210x.2012.00261.x

T. Snijders and R. Bosker, Modeled Variance in Two-Level Models, Sociological Methods & Research, vol.59, issue.3, pp.342-363, 1994.
DOI : 10.1177/0049124194022003004

F. Schoonjans, D. Bacquer, D. Schmid, and P. , Estimation of Population Percentiles, Epidemiology, vol.22, issue.5, pp.750-751, 2011.
DOI : 10.1097/EDE.0b013e318225c1de

D. Coomes and R. Allen, Effects of size, competition and altitude on tree growth, Journal of Ecology, vol.37, issue.5, pp.1084-1097, 2007.
DOI : 10.1139/x04-142

K. Kitajima and L. Poorter, Functional basis for resource niche partitioning by tropical trees, pp.172-188, 2008.

M. Walters, E. Kruger, and P. Reich, Growth, biomass distribution and CO2 exchange of northern hardwood seedlings in high and low light: relationships with successional status and shade tolerance, Oecologia, vol.33, issue.1, pp.7-16, 1993.
DOI : 10.1007/BF00317294

L. Poorter, F. Bongers, F. Sterck, and H. Wöll, ARCHITECTURE OF 53 RAIN FOREST TREE SPECIES DIFFERING IN ADULT STATURE AND SHADE TOLERANCE, Ecology, vol.84, issue.3, pp.602-608, 2003.
DOI : 10.2307/1938900

S. Thomas, Asymptotic Height as a Predictor of Growth and Allometric Characteristics in Malaysian Rain Forest Trees, American Journal of Botany, vol.83, issue.5, pp.556-566, 1996.
DOI : 10.2307/2445913

G. Koch, S. Sillett, G. Jennings, and S. Davis, The limits to tree height, Nature, vol.428, issue.6985, pp.851-854, 2004.
DOI : 10.1038/nature02417

H. Pretzsch and P. Biber, Size-symmetric versus size-asymmetric competition and growth partitioning among trees in forest stands along an ecological gradient in central Europe, Canadian Journal of Forest Research, vol.40, issue.2, pp.370-384, 2010.
DOI : 10.1139/X09-195

S. Gillner, N. Rüger, A. Roloff, and U. Berger, Low relative growth rates predict future mortality of common beech (Fagus sylvatica L.), Forest Ecology and Management, vol.302, pp.372-378, 2013.
DOI : 10.1016/j.foreco.2013.03.032

D. Johnson, D. Woodruff, K. Mcculloh, and F. Meinzer, Leaf hydraulic conductance, measured in situ, declines and recovers daily: leaf hydraulics, water potential and stomatal conductance in four temperate and three tropical tree species, Tree Physiology, vol.29, issue.7, pp.879-887, 2009.
DOI : 10.1093/treephys/tpp031

K. Kitajima and B. Bolker, Testing performance rank reversals among coexisting species: crossover point irradiance analysis, Functional Ecology, vol.130, issue.2, pp.276-281, 2003.
DOI : 10.1046/j.1469-8137.1999.00425.x

M. Wolf, Adaptive individual differences: the evolution of animal personalities [Doctoral dissertation] . The Netherland, 2009.

D. Tilman, Niche tradeoffs, neutrality, and community structure: A stochastic theory of resource competition, invasion, and community assembly, Proceedings of the National Academy of Sciences, vol.101, issue.30, pp.10854-10861, 2004.
DOI : 10.1073/pnas.0403458101

B. Beckage, L. Gross, W. Platt, W. Godsoe, and D. Simberloff, Individual variation and weak neutrality as determinants of forest diversity, Front Biogeogr, vol.3, pp.145-155, 2012.

J. Chase and J. Myers, Disentangling the importance of ecological niches from stochastic processes across scales, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.14, issue.3, pp.2351-2363, 2011.
DOI : 10.1111/j.1461-0248.2010.01584.x

W. Kendal, Taylor???s ecological power law as a consequence of scale invariant exponential dispersion models, Ecological Complexity, vol.1, issue.3, pp.193-209, 2004.
DOI : 10.1016/j.ecocom.2004.05.001

F. Valladares, D. Sanchez-gomez, and M. Zavala, Quantitative estimation of phenotypic plasticity: bridging the gap between the evolutionary concept and its ecological applications, Journal of Ecology, vol.84, issue.6, pp.1103-1116, 2006.
DOI : 10.1046/j.1365-3040.2002.00907.x